Boron

A primary function of boron is related to cell wall formation, so boron-deficient plants may be stunted. Sugar transport in plants, flower retention and pollen formation and germination also are affected by boron. Seed and grain production are reduced with low boron supply. Boron-deficiency symptoms first appear at the growing points. This results in a stunted appearance (rosetting), barren ears due to poor pollination, hollow stems and fruit (hollow heart) and brittle, discolored leaves and loss of fruiting bodies.

Copper

Copper is necessary for carbohydrate and nitrogen metabolism and, inadequate copper results in stunting of plants. Copper also is required for lignin synthesis which is needed for cell wall strength and prevention of wilting. Deficiency symptoms of copper are dieback of stems and twigs, yellowing of leaves, stunted growth and pale green leaves that wither easily.

Iron

Iron is involved in the production of chlorophyll, and iron chlorosis is easily recognized on iron-sensitive crops growing on calcareous soils. Iron also is a component of many enzymes associated with energy transfer, nitrogen reduction and fixation, and lignin formation. Iron is associated with sulfur in plants to form compounds that catalyze other reactions. Iron deficiencies are mainly manifested by yellow leaves due to low levels of chlorophyll. Leaf yellowing first appears on the younger upper leaves in interveinal tissues. Severe iron deficiencies cause leaves to turn completely yellow or almost white, and then brown as leaves die.

Manganese

Manganese is necessary in photosynthesis, nitrogen metabolism and to form other compounds required for plant metabolism. Interveinal chlorosis is a characteristic manganese-deficiency symptom. In very severe manganese cases, brown necrotic spots appear on leaves, resulting in premature leaf drop. Delayed maturity is another deficiency symptom in some species. White/gray spots on leaves of some cereal crops is a sign of manganese deficienc.