It is given in NCERT, that "mesophyll cells lack RuBP in C4 plants, but how can that be positive in C3 plants? 

Dear Student
In C4 plants mesophyll cell contains PEP carboxylase which helps in fixation of carbon whereas in C3 plants mesophyll cells contain Rubisco enzyme which fixes carbon. Mesophyll cells contain a majority of Rubisco enzyme in C3 Plants. 
Hope this information will clear your doubts about the topic.
Regards

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